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¸£Àû±ÆÕ¾ Catalog 2025-2026

Biological Sciences (BIO)

µþ±õ°¿Ìý105ÌýÌýBiology in the Modern WorldÌýÌý(3 credit hours)ÌýÌý

Concepts in biology taught from the perspective of civically engaging issues or problems. Themes and topics of wide interest, such as climate change, stem cells and cloning, antibiotic resistance, evolution, and human health. Lecture and the corresponding laboratory are designed to deepen an appreciation for the connections between science and "real-world" issues. For non-science majors. Students cannot receive credit for both µþ±õ°¿Ìý105 and (µþ±õ°¿Ìý181 or µþ±õ°¿Ìý183).

GEP Natural Sciences

Typically offered in Fall, Spring, and Summer

µþ±õ°¿Ìý106ÌýÌýBiology in the Modern World LaboratoryÌýÌý(1 credit hours)ÌýÌý

Laboratory experience in biological principles to complement µþ±õ°¿Ìý105. For non-science students. Students may not receive credit for both µþ±õ°¿Ìý106 and (µþ±õ°¿Ìý181 or µþ±õ°¿Ìý183).

Corequisite: µþ±õ°¿Ìý105; May not receive credit for both µþ±õ°¿Ìý106 and (µþ±õ°¿Ìý181 or µþ±õ°¿Ìý183)

GEP Natural Sciences

Typically offered in Fall, Spring, and Summer

µþ±õ°¿Ìý181ÌýÌýIntroductory Biology: Ecology, Evolution, and BiodiversityÌýÌý(4 credit hours)ÌýÌý

Introduction to ecology, evolution, and biodiversity within the context of structure and function including the adaptive strategies that organisms use to manage the everyday challenges of life. Emphasis on interactions of organisms with their environments, evolutionary change and role of natural selection in the evolution of life forms. Builds a foundation for understanding how science works, how to think critically and communicate scientifically.

X: Cannot receive credit for both µþ±õ°¿Ìý181 and µþ±õ°¿Ìý105 or µþ±õ°¿Ìý106.

GEP Natural Sciences

Typically offered in Fall, Spring, and Summer

µþ±õ°¿Ìý183ÌýÌýIntroductory Biology: Cellular and Molecular BiologyÌýÌý(4 credit hours)ÌýÌý

Basic concepts and principles of molecular, cellular, and developmental biology. Emphasis is placed on the physical basis of life, on the molecular mechanisms that guide evolution, on the cell as the fundamental unit of life, and on the mechanisms involved in the development of multicellular organisms. The course promotes the development of crucial scientific skills including critical thinking, problem solving, design of experiments, and effective oral and written scientific communication. Students cannot receive credit for both µþ±õ°¿Ìý183 and (µþ±õ°¿Ìý105 or µþ±õ°¿Ìý106).

GEP Natural Sciences

Typically offered in Fall, Spring, and Summer

µþ±õ°¿Ìý214ÌýÌýCellular and Molecular BiologyÌýÌý(3 credit hours)ÌýÌý

Exploration of cell biology with an emphasis on the integration of the chemistry of organic molecules in cellular function. Topics will include cell signaling, metabolism, and genetic regulation. Evolutionarily conserved mechanisms of action will be emphasized. Students should come to this course with an understanding of the concept of structure-function relationships in biology, the importance of chemical pathways in and between cells, and what it means for a gene, protein, or other structure to be evolutionarily conserved.

Prerequisite: µþ±õ°¿Ìý183

Typically offered in Fall only

µþ±õ°¿Ìý227ÌýÌýUnderstanding Structural Diversity through Biological IllustrationÌýÌý(3 credit hours)ÌýÌý

Biological concepts of diversity and anatomy taught through direct observation and illustrative techniques. Lecture topics include plant ID and structure, microscopic life forms, animal anatomy and identification. Laboratory work emphasizes close observation of structures and comparative anatomy as well as illustrative techniques to produce accurate drawings of specimens. Students will be required to provide their own transportation for one field trip.

GEP Interdisciplinary Perspectives, GEP Natural Sciences

Typically offered in Spring only

µþ±õ°¿Ìý230ÌýÌýThe Science of Studying DinosaursÌýÌý(3 credit hours)ÌýÌý

This course introduces students to the scientific method as applied to the study of dinosaurs and the world in which they lived. Because we cannot directly observe dinosaurs, estimating biological features such as running speed, growth rates, and reproductive and other physiological strategies presents challenges. We will examine a range of biological concepts (including cellular biology and physiology, functional morphology and biomechanics, evolutionary relationships, and paleoecology), as well as geological concepts (such as sedimentology, radiometric dating, plate tectonics, and the geologic time scale) as they apply to dinosaurs as living organisms. An understanding of biology at the introductory college level will be assumed.

Prerequisite: C- or better in µþ±õ°¿Ìý181 or µþ±õ°¿Ìý183 or µþ±õ°¿Ìý105

GEP Interdisciplinary Perspectives, GEP Natural Sciences

Typically offered in Fall only

µþ±õ°¿Ìý240ÌýÌýPrinciples of Human Anatomy & Physiology (A): Nervous, Skeletal, Muscular, & Digestive SystemsÌýÌý(3 credit hours)ÌýÌý

µþ±õ°¿Ìý240 provides an introduction to the anatomy and physiology of the nervous, skeletal, muscular, and digestive systems. Fundamental principles addressed throughout the course include (1) maintenance and regulation of homeostasis, (2) communication and control processes throughout the body, (3) integration and interdependence across organ systems, (4) structure-function relationships, and (5) anatomical and physiological adaptation. Students should come to this course with an understanding of the concept of structure-function relationships in biology, the importance of chemical pathways in and between cells, and what it means for a gene, protein, or other structure to be evolutionarily conserved.

Prerequisite: C- or better in µþ±õ°¿Ìý183

Typically offered in Fall, Spring, and Summer

µþ±õ°¿Ìý242ÌýÌýHuman Anatomy and Physiology LaboratoryÌýÌý(2 credit hours)ÌýÌý

µþ±õ°¿Ìý242 is a 2-credit hour laboratory course designed for undergraduate students preparing for a career in human health and others interested in human anatomy and physiology. This course focuses on scientific thinking with regard to the structures and functions of human body systems, with an emphasis on how these systems work together to maintain homeostasis. Students should come to this class with an understanding of 1) the close relationship between structure and function in biological systems, 2) the importance of chemical pathways in and between cells (e.g., metabolism, signaling), and 3) what it means for a gene, protein, or other structure to be evolutionarily conserved.

Prerequisite: C or better in µþ±õ°¿Ìý183; Corequisite: µþ±õ°¿Ìý240 or µþ±õ°¿Ìý245

Typically offered in Fall and Spring

µþ±õ°¿Ìý245ÌýÌýPrinciples of Human Anatomy & Physiology (B): Endocrine, Cardiovascular, Respiratory & Renal SystemsÌýÌý(3 credit hours)ÌýÌý

µþ±õ°¿Ìý245 provides an introduction to the anatomy and physiology of the endocrine, cardiovascular, respiratory, and renal systems. Fundamental principles addressed throughout the course include (1) maintenance and regulation of homeostasis, (2) communication and control processes throughout the body, (3) integration and interdependence across organ systems, (4) structure-function relationships, and (5) anatomical and physiological adaptation. Students should come to this course with an understanding of the concept of structure-function relationships in biology, the importance of chemical pathways in and between cells, and what it means for a gene, protein, or other structure to be evolutionarily conserved.

Prerequisite: C- or better in µþ±õ°¿Ìý183

Typically offered in Fall, Spring, and Summer

µþ±õ°¿Ìý267ÌýÌýResearch in the Life Sciences I: Research SkillsÌýÌý(3 credit hours)ÌýÌý

This course is designed to help first year students learn basic skills associated with scientific research. Class structure is interactive and relies on group collaboration for most projects. Students will become confident in reading and analyzing scientific literature, communicating scientific principles, compiling a poster presentation, presenting at scientific conferences, and attending local scientific symposia as well as practicing some basic laboratory techniques. The 2-semester Research PackTrack Program (µþ±õ°¿Ìý267 and 269) is designed to prepare undergraduates for an original research experience in a scientific laboratory. A B- or better in µþ±õ°¿Ìý267 is required to take µþ±õ°¿Ìý269. Students in µþ±õ°¿Ìý267 are required to attending one research symposium outside of regular class time. Individualized/Independent Study and Research courses require a Course Agreement for Students Enrolled in Non-Standard Courses be completed by the student and faculty member prior to registration by the department.

Prerequisite: C- or better in µþ±õ°¿Ìý181 and Corequisite: µþ±õ°¿Ìý183

Typically offered in Spring only

µþ±õ°¿Ìý269ÌýÌýResearch in the Life Sciences II: Guided ResearchÌýÌý(3 credit hours)ÌýÌý

This course is designed to provide students with a laboratory framework for conducting original research and (together with µþ±õ°¿Ìý267) preparation to move on to conducting research in a scientific laboratory. Students will generate their own research goals, write research proposals, conduct original mentored independent research, and present their findings at poster symposia both at NC State and at local conferences.

Prerequisite: C- or better in µþ±õ°¿Ìý183 and B- or better in µþ±õ°¿Ìý267

Typically offered in Fall and Spring

µþ±õ°¿Ìý270ÌýÌýIntroduction to EvolutionÌýÌý(3 credit hours)ÌýÌý

Evolution is the "grand unifying theory" that underlies all of modern biology. In this team-taught, lecture and discussion-based introductory course, students will learn about: the history of the concept of evolution and how it is discussed in modern society; sources of variation and forces of evolution; microevolution, speciation, and the nature of species; common descent, phylogeny, and the tree (and web) of life; macroevolution and modeling evolutionary trends; and the role of the fossil record, including that of our own lineage, in understanding evolution in deep time. Students will learn core foundational knowledge and how to discuss and contextualize evolutionary concepts.

Prerequisite: C- or better in µþ±õ°¿Ìý181 or in (µþ±õ°¿Ìý105 and 106)

Typically offered in Fall and Spring

µþ±õ°¿Ìý310ÌýÌýQuantitative Approaches to Biological ProblemsÌýÌý(3 credit hours)ÌýÌý

This course serves as an introduction to the use of mathematical, statistical and computational models and tools for understanding biology at the molecular, population, ecological and evolutionary scales. We will use discrete and continuous mathematics to model disease spread, ecological competition and biochemical systems. We explore the effect of random events in biochemistry and learn how to model such randomness in a statistical framework. We will also use computation tools to analyze genetic data and write a short computer program that simulates the change in allele frequencies in a population in time due to random effects. All course topics will involve hands-on computational exercises, but no prior experience in these tools and methods is expected.

Prerequisite: µþ±õ°¿Ìý181 and ²Ñ´¡Ìý131 or ²Ñ´¡Ìý141 (or equivalents).

Typically offered in Spring only

µþ±õ°¿Ìý315ÌýÌýGeneral ParasitologyÌýÌý(3 credit hours)ÌýÌý

General principles of parasitic symbiosis. Emphasis on life cycles, epidemiology, and pathology of major parasites of humans and domestic animals.

Prerequisite: C- or better in µþ±õ°¿Ìý181 and µþ±õ°¿Ìý183

Typically offered in Spring only

µþ±õ°¿Ìý323ÌýÌýPaleoecologyÌýÌý(3 credit hours)ÌýÌý

Paleoecology offers insights into the ancient ecosystems of Earth across millions of years and how the study of these extinct realms compares and contrasts with ecological studies of the modern world. Students will learn about ancient organisms through firsthand observation of fossil specimens, will have hands-on experience with paleontological data collection, and will learn to perform data analysis on paleontological datasets. At least one optional weekend field trip (illustrating concepts learned in class) will be offered during the semester.

Prerequisite: µþ±õ°¿Ìý181

Typically offered in Fall only

µþ±õ°¿Ìý325ÌýÌýPaleontological Field MethodsÌýÌý(4 credit hours)ÌýÌý

Understanding the history of life is not possible without first and foremost unearthing it. Discovering, collecting, and conserving fossil specimens for research is a time- and labor-intensive endeavor that requires hands-on training in the field. This course provides introductions to the evolution of life; Mesozoic biodiversity, biostratigraphy, and biogeography; vertebrate skeletal anatomy; stratigraphic and sedimentary concepts; and geomorphology. The course also covers paleontological research methods, including: specimen data collection, conservation, and preparation; topographic and geologic maps; GPS techniques; prospecting and excavation of fossil localities; and taphonomic indicators. Field work in Utah; additional costs involved.

Typically offered in Summer only

µþ±õ°¿Ìý330ÌýÌýEvolutionary BiologyÌýÌý(3 credit hours)ÌýÌý

Principles and patterns of organic evolution. Topics will include the origin of life, patterns of genetic variation, adaptations, natural selection, and the formation of species, the relationship between micro and macroevolution, and the importance of evolution to humans and medicine.

Prerequisite: C- or better in BIO 181and µþ±õ°¿Ìý183

Typically offered in Spring only

µþ±õ°¿Ìý361ÌýÌýDevelopmental BiologyÌýÌý(3 credit hours)ÌýÌý

In this course students will discover the amazing journey that cells must take to get from an egg to an embryo, form a mature adult, and reproduce in order to continue the life cycle. Students will relate science to everyday life using developmental biology as a forum to integrate many aspects of biology from the molecules in single cells to the complete organism and how it is influenced by evolution and the environment. Cannot receive credit for both µþ±õ°¿Ìý361 and ³Ò±·Ìý434.

Prerequisite: C- or better in µþ±õ°¿Ìý183

Typically offered in Fall only

µþ±õ°¿Ìý370ÌýÌýDevelopmental Anatomy of the VertebratesÌýÌý(3 credit hours)ÌýÌý

An integrated study of the functional anatomy, phylogeny, and embryonic development of organ systems in vertebrate animals.

Prerequisite: C- or better in µþ±õ°¿Ìý181 or BIO 140

Typically offered in Spring only

µþ±õ°¿Ìý375ÌýÌýDevelopmental Anatomy LaboratoryÌýÌý(2 credit hours)ÌýÌý

A hands-on study of embryonic development and organ systems in vertebrate animals, utilizing microscopic examination of living and preserved embryos, demonstrations of skeletons and mammalian organs, and dissections of preserved shark, salamander, and mink.

Prerequisite: C- or better in µþ±õ°¿Ìý181 or BIO 140

Typically offered in Spring only

µþ±õ°¿Ìý404/³§°ÕÌý404ÌýÌýEpidemiology and Statistics in Global Public HealthÌýÌý(3 credit hours)ÌýÌý

This course will provide a general introduction to the quantitative methods used in global health, combining elements of epidemiology and biostatistics. The course will focus on linear and logistic regression, survival analysis, traditional study designs, and modern study designs. Students will learn fundamental principles in epidemiology, including statistical approaches, and apply them to topics in global public health.

Prerequisite: B- or better in ³§°ÕÌý311 or ST/µþ±«³§Ìý350 or ³§°ÕÌý370 or ³§°ÕÌý371 and either GPH 201 or ³Ò°¿±áÌý201

Typically offered in Fall only

µþ±õ°¿Ìý405ÌýÌýFunctional HistologyÌýÌý(3 credit hours)ÌýÌý

Offered only as a distance education course via the internet. Functional Histology describes the cellular structure of tissues and organs. Human organs are emphasized, with brief consideration given to variation in other mammals. Tissue and organ structure is related to function, including examples of malfunction (histopathology). The course is especially appropriate for students planning a career in veterinary science, medicine, or allied health fields. Offered by distance education only.

Prerequisite: C- or better in µþ±õ°¿Ìý183

Typically offered in Summer only

µþ±õ°¿Ìý414ÌýÌýCell BiologyÌýÌý(3 credit hours)ÌýÌý

The chemical and physical bases of cellular structure and function with emphasis on methods and interpretations.

Prerequisite: C- or better in µþ±õ°¿Ìý183 and (°ä±áÌý221 or °ä±áÌý225)

Typically offered in Spring only

µþ±õ°¿Ìý416ÌýÌýCancer Cell BiologyÌýÌý(3 credit hours)ÌýÌý

Cancer cells are defective and yet they survive and often thrive despite having these defects. We will explore changes in tumor cells as they transition from normal cells - looking at disrupted growth signaling and cell death pathways, alterations in cell motility patterns and the induction of changes in other cells. Current research and treatments will be discussed. A focus will be on problem solving and critical thinking skills as we integrate case studies as well as primary and review literature into the course. This course will involve lecture, discussion, and group activities. Students will be expected to do assigned reading prior to coming to class, make presentations or lead discussions, analyze data, and design experiments.

P: C- or better in µþ±õ°¿Ìý183 and ³Ò±·Ìý311. JR or SR standing; Students may not earn credit for both BIO/PB 414 and µþ±õ°¿Ìý416.

Typically offered in Fall only

µþ±õ°¿Ìý418ÌýÌýCell Biology Research LabÌýÌý(2 credit hours)ÌýÌý

µþ±õ°¿Ìý418 is a laboratory course that emphasizes collaborative, inquiry-based learning through research. Students will design and implement experiments to explore mechanisms relating to cellular growth, metabolism, cell signaling pathways, environmental toxins, and/or intrinsic and extrinsic apoptotic processes in various cell lines. This course will include primary literature analysis, experimental design and implementation, maintenance of lab notebooks, data analysis, discussions, and presentations.

C: BIO/PB 414 or µþ±õ°¿Ìý416. JR or SR standing

Typically offered in Fall and Spring

µþ±õ°¿Ìý424ÌýÌýEndocrinologyÌýÌý(3 credit hours)ÌýÌý

This course will explore the function of hormones and bioactive compounds in regulating animal physiology and homeostasis. Topics will include a study of hormones and their mechanism of actions in regulating various biological processes including development and growth; reproduction; feeding, digestion and metabolism; ion and water balance; stress and immunity; and sex determination. The methods used to study hormones and their physiological functions will also be addressed. 80% of enrollment is restricted to Biological Sciences and Zoology students with the remaning 20% open for all other majors.

Prerequisite: C- or better in BIO 250 or BIO 212 or µþ±õ°¿Ìý240 or µþ±õ°¿Ìý245

Typically offered in Fall only

µþ±õ°¿Ìý432ÌýÌýEvolutionary MedicineÌýÌý(3 credit hours)ÌýÌý

This course introduces the rapidly emerging field of evolutionary medicine as one approach to appreciating evolution as a unifying principle of biology. The primary goal is for students to better understand the evolutionary nature of many issues connected to human health and to better understand the field of evolution via examples that are medically relevant. The course will require reading and discussing scientific literature. Credit is not allowed for both µþ±õ°¿Ìý432 and µþ±õ°¿Ìý330.

Prerequisite: Junior or Senior Standing and a C- or better in Bio 270 or ³Ò±·Ìý311

Typically offered in Spring only

µþ±õ°¿Ìý434ÌýÌýHormones and BehaviorÌýÌý(3 credit hours)ÌýÌý

This course will focus on the field of behavioral neuroendocrinology, which explores mechanisms by which hormones affect and are affected by behavior. The course will use highly interactive lectures and discussions of material from the primary literature (seminal papers and recent exciting contributions). Student participation in class discussions and in-class assignments will be critical components of the learning process.

Prerequisite:C- or better in BIO 212 or Bio 250 or (´¡±·³§Ìý205 & ´¡±·³§Ìý206) or µþ±õ°¿Ìý240 or µþ±õ°¿Ìý245

Typically offered in Spring only

µþ±õ°¿Ìý440ÌýÌýThe Human Animal: An Evolutionary PerspectiveÌýÌý(3 credit hours)ÌýÌý

An in-depth look at the evolution of a wide range of human behaviors, and some aspects of physiology as well. We will critically explore the perceptions we hold of ourselves and the research that has sought to lend new insights into the fundamental bases of human behavior. New uses of evolutionary theory, including the field of evolutionary psychology, will be examined using a comparative approach and careful readings from primary and secondary literature in evolutionary biology and psychology. Classes will be largely discussion based.

Typically offered in Spring only

µþ±õ°¿Ìý444ÌýÌýThe Biology of Love and SexÌýÌý(3 credit hours)ÌýÌý

The need to find and seduce a mate is one of the most powerful forces in biology. In this course, we will examine the biological factors that contribute to love and sex. We will adopt a broad evolutionarily-based perspective, examining a variety of strategies in both human and animal systems. Our readings and discussions will cover current hypotheses and experimental methodologies spanning genetics, neuroscience, and endocrinology.

P: C- or better in two of the following: µþ±õ°¿Ìý330, BIO 410, µþ±õ°¿Ìý424, µþ±õ°¿Ìý488, ³Ò±·Ìý311; R: Junior or Senior Standing

Typically offered in Spring only

µþ±õ°¿Ìý481ÌýÌýSenior Capstone ProjectÌýÌý(1 credit hours)ÌýÌý

This course provides students an opportunity to integrate and apply knowledge and skills gained from their major studies in addressing an important challenge or problem that they identify. Emphasis will be placed on (1) reflections on the value of experiences outside of the classroom, (2) using discipline-specific knowledge and approaches from both biology and from the second disciplinary area that they chose to study within the B.A. in Biological Sciences, and (3) peer- and self-critiques based on the intellectual standards of critical and creative thinking. Restricted to seniors who will graduate with a B.A. in Biological Sciences. 17BIOBA majors only.

Typically offered in Fall and Spring

µþ±õ°¿Ìý482ÌýÌýCapstone Course in Molecular, Cellular, and Developmental BiologyÌýÌý(3 credit hours)ÌýÌý

Topical problems in molecular, cellular, and developmental biology. µþ±õ°¿Ìý482 provides a challenging opportunity for students to integrate and apply knowledge and skills gained from their major studies. Emphasis will be placed on collaborative learning and on effective, professional communication. Topics and instructors will vary from semester to semester. Priority will initially be given to seniors in the MCD curriculum; other students with the necessary prerequisites will be admitted on a space available basis.

Prerequisite: C- or better in µþ±õ°¿Ìý361, BIO/PB 414, and one of the following: µþ°ä±áÌý351 or µþ°ä±áÌý451 or µþ±õ°ÕÌý410 or ³Ò±·Ìý311.

Typically offered in Fall and Spring

µþ±õ°¿Ìý483ÌýÌýCapstone Course in Integrative Physiology and NeurobiologyÌýÌý(3 credit hours)ÌýÌý

Topical problems in integrative physiology and neurobiology. µþ±õ°¿Ìý483 provides a challenging opportunity for students to integrate and apply knowledge and skills gained from their major studies. Emphasis will be placed on collaborative learning and on effective, professional communication. Topics and instructors will vary from semester to semester, and the course description for each topic will indicate what prior knowledge is important for students to have for that offering.

Typically offered in Fall and Spring

µþ±õ°¿Ìý484ÌýÌýCapstone Course in Human BiologyÌýÌý(3 credit hours)ÌýÌý

Topical problems in human biology. µþ±õ°¿Ìý484 provides a challenging opportunity for students to integrate and apply knowledge and skills gained from their major studies. Emphasis will be placed on collaborative learning and on effective, professional communication. Topics and instructors will vary from section to section, and the course description for each topic will indicate what prior knowledge is important for students to have for that offering.

Typically offered in Fall and Spring

µþ±õ°¿Ìý485ÌýÌýCapstone Course in Ecology, Evolution, and Conservation BiologyÌýÌý(3 credit hours)ÌýÌý

Topical problems in ecology, evolution, and conservation biology. µþ±õ°¿Ìý485 provides a challenging opportunity for students to integrate and apply knowledge and skills gained from their major studies. Emphasis will be placed on collaborative learning and on effective, professional communication. Topics and instructors will vary from semester to semester. Priority will initially be given to seniors in the EEC curriculum; other students with the necessary prerequisites will be admitted on a space available basis.

Prerequisite: C- or better in BIO/PB 330, BIO/±ÊµþÌý360, and one of the following: BIO 460 or ³Ò±·Ìý311 or ±·¸éÌý406 or ³§°ÕÌý311.

Typically offered in Fall and Spring

µþ±õ°¿Ìý488/µþ±õ°¿Ìý588ÌýÌýNeurobiologyÌýÌý(3 credit hours)ÌýÌý

Overview of the neurosciences, with a focus on fundamental principles in the function, structure, and development of the nervous system. Topics include neuroanatomy, electrical signaling, synaptic transmission, sensory and motor systems, neural development, neural plasticity, and complex brain functions. Multiple levels of analysis, from molecular to behavioral, with an emphasis on the mammalian nervous system. Motivated students who do not meet listed prerequisite can contact the instructor for permission to take the course.

Prerequisite: C- or better in BIO 250 or BIO 212 or µþ±õ°¿Ìý240 or µþ±õ°¿Ìý245 or permission of instructor

Typically offered in Fall only

µþ±õ°¿Ìý520ÌýÌýSkeletal Biological Laboratory Methods in Human Identification & Cold CasesÌýÌý(3 credit hours)ÌýÌý

This laboratory-based course covers skeletal biological methods such as those used in human identification applied to medicolegal issues or issues of the law. Forensic identification methods will be introduced via lecture and students will apply these methods in the laboratory. The investigation of cold cases will be addressed through laboratory methodologies that intersects with missing persons cases. This course covers a broad array of investigative skeletal methods. However, students will not be qualified to practice after taking this course.

P: Graduate Standing

Typically offered in Fall only

µþ±õ°¿Ìý555/±Ê³§°äÌý555ÌýÌýCreative Media Production for ScientistsÌýÌý(3 credit hours)ÌýÌý

This course will be an introduction to producing videos and other online science media. Students will survey and discuss research-based best practices for online science media and will produce their own media pieces.

R: Grad Standing or Instructor Approval

Typically offered in Spring only

µþ±õ°¿Ìý560/µþ²Ñ´¡Ìý560ÌýÌýPopulation EcologyÌýÌý(3 credit hours)ÌýÌý

Dynamics of natural populations. Current work, theories and problems dealing with population growth, fluctuation, limitation and patterns of dispersion, species interactions, community structure and ecological genetics. One semester of calculus and a junior/senior level ecology course are required.

Co-requisite: ³§°ÕÌý511

Typically offered in Spring only

This course is offered alternate odd years

µþ±õ°¿Ìý562ÌýÌýFundamentals of BioinformaticsÌýÌý(3 credit hours)ÌýÌý

Almost every aspect of modern biology involves large-scale datasets and computational analyses. This course will cover some of the basic theoretical and practice background needed to understand and use computational tools for biological analyses. The course will feature a mixture of lecture, activity-based and hand-on computational analyses using the LINUX operating system. A background in biology or computer science is recommended.

Typically offered in Fall only

µþ±õ°¿Ìý570ÌýÌýEvolutionary EcologyÌýÌý(3 credit hours)ÌýÌý

This course provides a detailed overview of evolutionary ecology, the multidisciplinary interface of ecological and evolutionary processes. The course includes an historical perspective of the development of the field, major theoretical and empirical milestones, and the cutting edge of modern evolutionary ecology research and application. This is a discussion-oriented course, drawing heavily from peer-reviewed literature. Grading is centered on participation, leading discussions, critical literature reviews, and a research project. The course Moodle website will contain files for all assigned readings, the course syllabus, tips for leading discussions, and a schedule of class meetings and discussion leaders.

Prequisite: Graduate Standing or C- or better in AEC/±ÊµþÌý360 and Corequisite: C- or better in µþ±õ°¿Ìý330 or µþ±õ°¿Ìý432

Typically offered in Fall only

µþ±õ°¿Ìý572/°ä±áÌý572/µþ±õ°ÕÌý572ÌýÌýProteomicsÌýÌý(3 credit hours)ÌýÌý

Introduction and history of the field of proteomics followed by the principles and applications of proteomics technology to understand protein expression and protein post-transitional modifications. Laboratory sessions include growing yeast with stable-isotope labeled amino acids, protein purification, Western blots, protein identification and quantification, and protein bioinformatic analysis. This is a half-semester course.

Prerequisite: µþ±õ°ÕÌý410 or µþ±õ°ÕÌý510 or µþ°ä±áÌý454 (or approval from the instructor)

Typically offered in Spring only

µþ±õ°¿Ìý578ÌýÌýThe Physiology of StressÌýÌý(3 credit hours)ÌýÌý

Stress is a nearly unavoidable component of modern life, bound to affect each of us at some point during the week (or day!) And stress can wreak havoc on both mental and physical health. Beginning with a look at the normal stress response in humans and animals, this course then examines the biological effects of both physical and psychological stress. We'll examine what happens to various systems in the body when the stress response occurs and doesn't shut down. Among the systems explored will be neuroendocrine, cardiovascular, digestive, immune, and reproductive. We'll also look at how stress affects sleep, depression, prenatal and childhood development, and memory and judgment. And we'll spend some time with stress management techniques (and why they do or don't work) as well.

P: Graduate Standing

Typically offered in Spring only

µþ±õ°¿Ìý588/µþ±õ°¿Ìý488ÌýÌýNeurobiologyÌýÌý(3 credit hours)ÌýÌý

Overview of the neurosciences, with a focus on fundamental principles in the function, structure, and development of the nervous system. Topics include neuroanatomy, electrical signaling, synaptic transmission, sensory and motor systems, neural development, neural plasticity, and complex brain functions. Multiple levels of analysis, from molecular to behavioral, with an emphasis on the mammalian nervous system. Motivated students who do not meet listed prerequisite can contact the instructor for permission to take the course.

Prerequisite: C- or better in BIO 250 or BIO 212 or µþ±õ°¿Ìý240 or µþ±õ°¿Ìý245 or permission of instructor

Typically offered in Fall and Summer

µþ±õ°¿Ìý592ÌýÌýTopical ProblemsÌýÌý(1-3 credit hours)ÌýÌý

Organized, formal lectures and discussion of a special topic.

Typically offered in Fall, Spring, and Summer

µþ±õ°¿Ìý624ÌýÌýTopical ProblemsÌýÌý(1-3 credit hours)ÌýÌý

Organized, formal lectures and discussion of a special topic.

Typically offered in Fall and Spring

µþ±õ°¿Ìý705/°äµþ³§Ìý705ÌýÌýFundamentals of NeuroscienceÌýÌý(3 credit hours)ÌýÌý

This is a fundamental course that will provide the student with an up-to-date coverage of molecular, cellular, physiological, and circuit-based aspects of Modern Neurosciences in the Comparative Biomedical Sciences Graduate Program. Being a graduate-level class, the instructors will assume that the students have acquired some background in basic biology and biochemistry. The most important goal of this course is to train PhD students in neuroscience function and disorders, preparing them for neuroscience research. Emphasis will be placed on the systems and skills needed to pursue experimental neuroscience activities. Important components of the learning process will be presentations from neuroscience experts, class discussions, exams and in class activities.

P: µþ±õ°¿Ìý488 or permission of instructor

Typically offered in Fall only

µþ±õ°¿Ìý727/°ä±áÌý727ÌýÌýBiological Mass SpectrometryÌýÌý(3 credit hours)ÌýÌý

Fundamentals of mass spectrometry including topics such as: mass, isotopic distributions, resolving power, mass accuracy. Ionization source topics: electron impact, chemical ionization, matrix-assisted laser desorption ionization, electrospray ionization and contemporary methods. Instrumentation and mass analyzers: quadrupole, time-of-flight, Fourier transform based mass analyzers; hybrid instruments such as a quadrupole orbitrap. Tandem mass spectrometry and dissociation. Applications: quantitation, small molecule analysis, and peptide sequencing.

Typically offered in Fall only

This course is offered alternate years

µþ±õ°¿Ìý792ÌýÌýTopical ProblemsÌýÌý(1-6 credit hours)ÌýÌý

Organized, formal lectures and discussion of a special topic.

Typically offered in Fall, Spring, and Summer

µþ±õ°¿Ìý805/°äµþ³§Ìý805ÌýÌýSpecial Topics in NeuroscienceÌýÌý(1 credit hours)ÌýÌý

Topics in neuroscience. This course will provide an opportunity for students to integrate and apply knowledge and skills gained from their graduate studies. Emphasis will be placed on primary literature, scientific practice, and on effective, professional communication and presentations. Topics and instructors will vary from semester to semester. Priority will initially be given to graduate students participating in the neuroscience concentration; other students with the necessary prerequisites will be admitted on a space available basis.

R: Grad Standing or Instructor Approval

Typically offered in Spring only

µþ±õ°¿Ìý811ÌýÌýForensic Sciences SeminarÌýÌý(1 credit hours)ÌýÌý

This is a webinar series that includes professional development as well as practitioner presentations about the current state of the various forensic sciences disciplines. Topics covered fill in the background and gaps needed for a career as a forensic science practitioner and/or academician.

Prerequisite: Graduate Standing or Instructor Permission

Typically offered in Fall only

µþ±õ°¿Ìý824ÌýÌýTopical ProblemsÌýÌý(1-3 credit hours)ÌýÌý

Organized, formal lectures and discussion of a special topic.

Typically offered in Fall and Spring